List of European animals extinct in the Holocene
This list of European animals extinct in the Holocene features animals known to have become extinct in the last 12,000 years on the European continent and its surrounding islands.

All large islands in the Mediterranean Sea are included except for Cyprus, which is in the List of Asian animals extinct in the Holocene. The recently extinct animals of the Macaronesian islands in the North Atlantic are listed separately. Overseas territories of European countries are not included here; they are found on the lists pertaining to their respective regions. For example, French Polynesia is grouped with Oceania, while Réunion is grouped with Madagascar and the Indian Ocean islands.
Many extinction dates are unknown due to a lack of relevant information.
Mammals
Undated
| Common name/scientific name | Range |
|---|---|
| Ochotona transcaucasica | the Caucasus[1] |
Prehistoric
| Common name/scientific name | Extinction date | Range | Image |
|---|---|---|---|
| Sardinian giant shrew Asoriculus similis |
3050 BCE[2] | Sardinia, Italy | |
| Steppe bison Bison priscus |
1130-1060 BCE[3] | Northern Eurasia and North America | ![]() |
| Woolly rhinoceros Coelodonta antiquitatis |
7820-7300 BCE[4] | Northern Eurasia | ![]() |
| European dhole Cuon alpinus europaeus |
7050-6550 BCE[5] | Central and Southern Europe; the Caucasus?[6] | ![]() |
| Sardinian dhole Cynotherium sardous |
9500-9300 BCE[7] | Corsica and Sardinia | ![]() |
| European wild ass Equus hemionus hydruntinus |
3200-2500 BCE[8] | Europe and Southwest Asia | ![]() |
| Majorcan giant dormouse Hypnomys morpheus |
4840-4690 BCE[9] | the Gymnesian Islands, Spain | ![]() |
| Woolly mammoth Mammuthus primigenius |
9290-8970 BCE[4] | Northern Eurasia and North America | ![]() |
| Irish elk Megaloceros giganteus |
4912-4846 BCE[3] | Europe and Southern Siberia | ![]() |
| Balearic Islands cave goat Myotragus balearicus |
2830-2470 BCE[10] | the Gymnesian Islands, Spain | ![]() |
| Balearic giant shrew Nesiotites hidalgo |
3030-2690 BCE[9] | the Gymnesian Islands, Spain | ![]() |
| Tilos dwarf elephant Palaeoloxodon tiliensis |
3040-1840 BCE[11] | Tilos, Greece | ![]() |
| Sardinian deer Praemegaceros cazioti |
5550 BCE[12] | Corsica and Sardinia[13] | ![]() |
| Sardinian pika Prolagus sardus |
348 BCE - 283 CE[14] | Corsica and Sardinia | ![]() |
| Tyrrhenian field rat Rhagamys orthodon |
348 BCE - 283 CE[14] | Corsica and Sardinia | ![]() |
| Tyrrhenian vole Tyrrhenicola henseli |
348 BCE - 283 CE[14] | Corsica and Sardinia |
Recent
| Common name/scientific name | Extinction date | Range | Image |
|---|---|---|---|
| Caucasian elk Alces alces caucasicus |
c. 1900[15] | Northern Caucasus and the Transcaucasian coast of the Black Sea | ![]() |
| Caucasian wisent Bison bonasus caucasicus |
1927[16] | the Caucasus Mountains | ![]() |
| Eurasian aurochs Bos primigenius primigenius |
1627[17] | Mid-latitude Eurasia | ![]() |
| Sicilian wolf Canis lupus cristaldii |
1970[18] | Sicily, Italy | ![]() |
| Portuguese ibex Capra pyrenaica lusitanica |
c. 1890[19] | the Portuguese-Galician border | ![]() |
| Pyrenean ibex Capra pyrenaica pyrenaica |
2000[upper-alpha 1] |
the Pyrenees and possibly the Cantabrian Mountains[21] | ![]() |
| Tarpan Equus ferus ferus |
1909[22] | Europe | ![]() |
| St. Kilda house mouse Mus musculus muralis |
1930[23] | St. Kilda, Scotland | ![]() |
| Caspian tiger Panthera tigris virgata |
1922[24] | the Caucasus, Western and Central Asia | ![]() |
Local
| Common name/scientific name | Extinction date | Range | Reintroduction | Image |
|---|---|---|---|---|
| Lowland wisent Bison bonasus bonasus |
1919[25] | Central Europe to southern Siberia | 1946[26] | ![]() |
| Wapiti Cervus canadensis |
Early Holocene[27] | Central and Northern Europe to Asia and North America | ![]() | |
| Turkmenian kulan Equus hemionus kulan |
18th-19th century[28] | Ukraine to Central Asia[29] | 1950[30] | ![]() |
| Gray whale Eschrichtius robustus |
550[31] | the North Atlantic, the Mediterranean Sea, and the northern Pacific Ocean[32] | 2010[upper-alpha 2] | |
| Barbary macaque Macaca sylvanus |
c. 70,000-60,000 BCE[35] | Central and Southern Europe and North Africa | 1704[36] | ![]() |
| Narrow-headed vole Microtus gregalis |
Late Holocene[37][38] | Northern Eurasia | ![]() | |
| Steppe pika Ochotona pusilla |
Late Holocene[37][38] | Western Europe to Kazakhstan | ![]() | |
| Muskox Ovibos moschatus |
7050 BCE[39] | Northern Eurasia and North America | 1947[40][upper-alpha 3] | ![]() |
| Lion Panthera leo |
4th century (Balkans)[41] 10th century (Caucasus)[42] |
Africa, the Middle East, northern India, and southeastern Europe | ![]() |
Birds
| Common name/scientific name | Extinction date | Range |
|---|---|---|
| Mediterranean brown fish owl Ketupa zeylonensis lamarmorae |
7433-7035 BCE[14] | Corsica, Sardinia, and Crete[43] |
| Ibiza rail Rallus eivissensis |
5295-4848 BCE[44] | Ibiza, Spain |
| Common name/scientific name | Extinction date | Range | Image |
|---|---|---|---|
| Great auk Pinguinus impennis |
1844[45] | the northern Atlantic and the western Mediterranean | ![]() |
| Pied raven Corvus corax varius morpha leucophaeus |
1902[46] | the Faroe Islands, Denmark | ![]() |
| Common name/scientific name | Extinction date | Range | Image |
|---|---|---|---|
| Slender-billed curlew Numenius tenuirostris |
2001[47] | Western Eurasia and North Africa | ![]() |
| Common name/scientific name | Extinction date | Range | Reintroduction | Image |
|---|---|---|---|---|
| Marsh owl Asio capensis |
1998[48] | Africa and the southwestern Iberian Peninsula | ![]() | |
| Northern bald ibis Geronticus eremita |
16th century | the Mediterranean region | 2004[49] | ![]() |
| Common buttonquail Turnix sylvaticus |
1981[50] | Africa, South Asia, the southwestern Iberian Peninsula, and Sicily | ![]() |
Reptiles
| Common name/scientific name | Extinction date | Range | Image |
|---|---|---|---|
| Ratas Island lizard Podarcis lilfordi rodriquezi |
1935[51] | Ratas Island off Mahón, Spain | ![]() |
| Santo Stefano lizard Podarcis sicula sanctistephani |
c. 1965[52] | Santo Stefano Island, Italy |
Fish
| Common name/scientific name | Extinction date | Range | Image |
|---|---|---|---|
| Skadar nase Chondrostoma scodrense |
1900s[53] | Lake Skadar | |
| Coregonus bezola | 1960s[54] | Lac du Bourget, France | |
| Coregonus fera | 1920[55] | Lake Geneva | |
| Lake Constance whitefish Coregonus gutturosus |
early 1970s[56] | Lake Constance | ![]() |
| Gravenche Coregonus hiemalis |
1950[57] | Lake Geneva | ![]() |
| Houting Coregonus oxyrinchus |
1940[58] | the southern North Sea, the Scheldt, Meuse and Rhine Basin up to Cologne, and southeastern England | ![]() |
| Coregonus restrictus | 1890[59] | Lake Morat, Switzerland | |
| Ukrainian migratory lamprey Eudontomyzon sp. nov. 'migratory' |
before 1900[60] | the Dniestr, Dniepr, and Don River drainages | |
| Techirghiol stickleback Gasterosteus crenobiontus |
1960s[61] | Lake Techirghiol, Romania | |
| Danube delta gudgeon Romanogobio antipai |
1960s[62] | the Lower Danube | |
| Salvelinus neocomensis | 1904[63] | Lake Neuchâtel, Switzerland |
| Common name/scientific name | Extinction date | Range | Image |
|---|---|---|---|
| Beloribitsa Stenodus leucichthys |
1960s[64] | the Caspian Sea, the Volga, Ural and Terek River drainages |
Insects
| Common name/scientific name | Range |
|---|---|
| Perrin's cave beetle Siettitia balsetensis |
France[65] |
| Common name/scientific name | Extinction date | Range |
|---|---|---|
| Tobias' caddisfly Hydropsyche tobiasi |
1938[66] | the Rhine and Main River, Germany |
| British large copper Lycaena dispar dispar |
1864[67] | the British Isles |
| Moss-land silver-studded blue Plebejus argus masseyi |
1942[68] | Lancashire and Cumbria, the United Kingdom |
| Pseudoyersinia brevipennis | 1860[69] | Hyères, France |
| Common name/scientific name | Extinction date | Range |
|---|---|---|
| Iberian lynx louse Felicola isidoroi |
1997[70] | the Iberian Peninsula |
Sea anemones
| Common name/scientific name | Extinction date | Range | Image |
|---|---|---|---|
| Ivell's sea anemone Edwardsia ivelli |
1983[71] | the Widewater Lagoon, West Sussex, United Kingdom | ![]() |
Molluscs
| Common name/scientific name | Range |
|---|---|
| Zonites siphnicus | Sifnos, Sikinos, and Folegandros, Greece[72] |
| Common name/scientific name | Extinction date | Range |
|---|---|---|
| Zonites santoriniensis | c. 1600 BCE[73] | Santorini, Greece |
| Common name/scientific name | Extinction date | Range | Image |
|---|---|---|---|
| Graecoanatolica macedonica | 1988[74] | Doiran Lake, the Greece-North Macedonia border | ![]() |
| Ohridohauffenia drimica | before 1983[75] | the Drin River, North Macedonia |
| Common name/scientific name | Extinction date | Range |
|---|---|---|
| Belgrandia varica | 1910[76] | the Var River Delta, France |
| Belgrandiella boetersi | unknown[77] | Tiefsteinschlucht, Austria |
| Parmacella gervaisii | 1874[78] | La Crau, Provence, France |
| Zonites embolium | 1985[79] | Zaforas, Greece |
See also
- Limousin horse, extinct
- List of extinct animals of Catalonia
- List of extinct animals of Caucasus
- List of extinct animals of the British Isles
- List of extinct and endangered species of Italy
- List of extinct and endangered animals of Lithuania
- List of extinct animals of the Netherlands
- List of extinct species
- List of extinct bird species since 1500
- Extinct in the wild
- Lazarus taxon
Notes
References
- Averianov, A. (2001). Pleistocene lagomorphs of Eurasia. Deinsea, 8(1), 1-14.
- Louys, J.; Braje, T. J.; Chang, C.-H.; Cosgrove, R.; Fitzpatrick, S. M.; Fujita, M.; Hawkins, S.; Ingicco, T.; Kawamura, A.; MacPhee, R. D. E.; McDowell, M. C.; Meijer, H. J. M.; Piper, P. J.; Roberts, P.; Simmons, A. H.; van den Bergh, G.; van der Geer, A.; Kealy, S.; O'Connor, S. (2021). "No evidence for widespread island extinctions after Pleistocene hominin arrival". Proceedings of the National Academy of Sciences of the United States of America. 118 (20): e2023005118. Bibcode:2021PNAS..11823005L. doi:10.1073/pnas.2023005118. ISSN 0027-8424. PMC 8157961. PMID 33941645.
- Plasteeva, N. A., Gasilin, V. V., Devjashin, M. M., & Kosintsev, P. A. (2020). Holocene Distribution and Extinction of Ungulates in Northern Eurasia. Biology Bulletin, 47(8), 981-995.
- Kosintsev, P. (2007). Late Pleistocene large mammal faunas from the Urals. Quaternary International, 160(1), 112-120.
- Ripoll, M. P., Perez, J. V. M., Serra, A. S., Tortosa, J. E. A., & Montanana, I. S. (2010). Presence of the genus Cuon in upper Pleistocene and initial Holocene sites of the Iberian Peninsula: new remains identified in archaeological contexts of the Mediterranean region. Journal of Archaeological Science, 37(3), 437-450.
- Ghezzo, E., & Rook, L. (2014). Cuon alpinus (Pallas, 1811)(Mammalia, Carnivora) from Equi (Late Pleistocene, Massa-Carrara, Italy): anatomical analysis and palaeoethological contextualisation. Rendiconti Lincei, 25(4), 491-504.
- Turvey, Sam (2009). Holocene extinctions. Oxford University Press. ISBN 978-0-19-953509-5. Retrieved 29 February 2012.
- Crees, Jennifer J.; Turvey, Samuel T. (May 2014). "Holocene extinction dynamics of Equus hydruntinus, a late-surviving European megafaunal mammal". Quaternary Science Reviews. 91: 16–29.
- Bover, P. (2011). La paleontologia de vertebrats insulars de les Balears: la contribució de les excavacions recents. Endins: publicació d'espeleologia, 299-316.
- Bover, P., et al. (2016). Closing the gap: new data on the last documented Myotragus and the first human evidence on Mallorca (Balearic Islands, Western Mediterranean Sea). The Holocene, 26(11), 1887-1891.
- Masseti, M. (2008). The most ancient explorations of the Mediterranean. Proc. Calif. Acad. Sci. 4th Ser, 59(Suppl I), 1-18.
- Benzi, V. et al. (2007). Radiocarbon and U-series dating of the endemic deer Praemegaceros cazioti (Depéret) from "Grotta Juntu", Sardinia. Journal of archaeological science, 34(5), 790-794.
- Melis, S., Salvadori, S., & Pillola, G. L. (2010). SARDINIAN DEER: DERIVATIONS, FOSSIL DISCOVERIES AND CURRENT DISTRIBUTION. Present Environment & Sustainable Development, 4(2).
- Vigne, Jean-Denis, Salvador Bailon, and Jacques Cuisin. "Biostratigraphy of amphibians, reptiles, birds and mammals in Corsica and the role of man in the Holocene faunal turnover." Anthropologica 25.26 (1997): 587-604.
- Boeskorov, G.G. (2003) The genetics of the modern moose and a review of its taxonomy. Cranium 20, Vol. 2: 31-45.
- Bashkirov, I. S. (1939). "Caucasian European Bison". Moscow: Central Board for Reserves, Forest Parks and Zoological Gardens, Council of the People's Commissars of the RSFSR: 1–72. [In Russian.]
- Van Vuure, C., & van Vuure, T. (2005). Retracing the aurochs: history, morphology and ecology of an extinct wild ox. Pensoft Pub.
- Angelici, F. M.; Rossi, L. (2018). "A new subspecies of grey wolf (Carnivora, Canidae), recently extinct, from Sicily, Italy" (PDF). Bollettino del Museo Civico di Storia Naturale di Verona. 42: 3–15.
- Acevedo, P., & Cassinello, J. (2009). Biology, ecology and status of Iberian ibex Capra pyrenaica: a critical review and research prospectus. Mammal Review, 39(1), 17-32.
- J. Folch; J. Cocero; M. J. Chesne; P. Alabart; J. K. Dominguez; V. Congnie; Y. Roche; A. Fernández-Árias; A. Marti; J. I. Sánchez; P. Echegoyen; E. Beckers; J. F. Sánchez; A. Bonastre; X. Vignon (2009). "First birth of an animal from an extinct subspecies (Capra pyrenaica pyrenaica) by cloning". Theriogenology. 71 (#6): 1026–1034. doi:10.1016/j.theriogenology.2008.11.005. PMID 19167744.
- Ríu, J. U. (1959). El "mueyu", "capra pyrenaica" asturiana extinguida a comienzos del siglo pasado. Archivum: Revista de la Facultad de Filología, (9), 361-375.
- Tadeusz Jezierski, Zbigniew Jaworski: Das Polnische Konik. Die Neue Brehm-Bücherei Bd. 658, Westarp Wissenschaften, Hohenwarsleben 2008
- "People and nature on St Kilda". www.ihbc.org.uk. Archived from the original on 13 March 2016. Retrieved 12 January 2022.
- Rossi, L., Scuzzarella, C. M., & Angelici, F. M. (2020). Extinct or Perhaps Surviving Relict Populations of Big Cats: Their Controversial Stories and Implications for Conservation. In Problematic Wildlife II (pp. 393-417). Springer, Cham.
- Plumb, G.; Kowalczyk, R.; Hernandez-Blanco, J.A. (2020). "Bison bonasus". IUCN Red List of Threatened Species. 2020: e.T2814A45156279. doi:10.2305/IUCN.UK.2020-3.RLTS.T2814A45156279.en. Retrieved 19 November 2021.
- Sipko, T. P. (2009). European bison in Russia–past, present and future. European Bison Conservation Newsletter, 2, 148-159.
- Croitor, R. (2020). A new form of wapiti Cervus canadensis Erxleben, 1777 (Cervidae, Mammalia) from the Late Pleistocene of France. Palaeoworld, 29(4), 789-806.
- Heptner, V. G., Nasimovich, A. A., Bannikov, A. G., & Hoffman, R. S. (1989). Mammals of the Soviet Union, vol. 1. Leiden, the Netherlands: EJ Brill, 1147 pages.
- Kaczensky, P., Lkhagvasuren, B., Pereladova, O., Hemami, M., & Bouskila, A. (2015). Equus hemionus. The IUCN red list of threatened species 2015: e. T7951A45171204.
- Yasinetskaya, N.I. (1997) НАУЧНОЕ И ЭКОЛОГО-ПРОСВЕТИТЕЛЬСКОЕ ЗНАЧЕНИЕ КОЛЛЕКЦИИ ПРЕДСТАВИТЕЛЕЙ СЕМЕЙСТВА ЛОШАДИНЫХ EQUIDAE ЗООПАРКА "АСКАНИЯ-НОВА". In Современные проблемы зоологии, экологии и охраны природы. Материалы чтений и научной конференции, посвященных памяти профессора Андрея Григорьевича Банникова, и 100-летию со дня его рождения. ЕВРОАЗИАТСКАЯ РЕГИОНАЛЬНАЯ АССОЦИАЦИЯ ЗООПАРКОВ И АКВАРИУМОВ, 351 pages.
- Jones, M.L. et al. (2012) The Gray Whale: Eschrichtius robustus. Academic Press, 600 pages.
- IUCN.
- Hamilton, Alex (October 8, 2015). "The Gray Whale Sneaks Back into the Atlantic, Two Centuries Later". WNYC. Retrieved 2019-10-28.
- Schiffman, Richard (February 25, 2016). "Why Are Gray Whales Moving to the Ocean Next Door?". Discover Magazine. Retrieved 2019-10-28.
- Rosendahl, W., Ambros, D., Hilpert, B., Hambach, U., Alt, K. W., Knipping, M., ... & Kaulich, B. (2011). Neanderthals and monkeys in the Würmian of Central Europe: the middle Paleolithic site of Hunas, southern Germany. In Neanderthal lifeways, subsistence and technology (pp. 15-23). Springer, Dordrecht.
- Modolo, L., Salzburger, W., & Martin, R. D. (2005). Phylogeography of Barbary macaques (Macaca sylvanus) and the origin of the Gibraltar colony. Proceedings of the National Academy of Sciences, 102(20), 7392-7397.
- Németh, A., Bárány, A., Csorba, G., Magyari, E., Pazonyi, P., & Pálfy, J. (2017). Holocene mammal extinctions in the Carpathian Basin: a review. Mammal Review, 47(1), 38-52.
- Kosintsev, P. A., & Bachura, O. P. (2014). Formation of recent ranges of mammals in the Urals during the Holocene. Biology Bulletin, 41(7), 629-637.
- Peter C. Lent (1999). Muskoxen and Their Hunters: A History. University of Oklahoma Press. ISBN 978-0-8061-3170-2. Retrieved 2013-08-25.
- Jorgensen, D. (2019) Recovering lost species in the modern age: Histories of longing and belonging. MIT Press, 256 pages.
- Hughes, J. D. (2003). Europe as consumer of exotic biodiversity: Greek and Roman times. Landscape Research, 28(1), 21-31.
- Heptner, V. G.; Sludskiy, A. A. (1992) [1972]. "Lion". Mlekopitajuščie Sovetskogo Soiuza. Moskva: Vysšaia Škola [Mammals of the Soviet Union, Volume II, Part 2]. Washington DC: Smithsonian Institution and the National Science Foundation. pp. 83–95. ISBN 978-90-04-08876-4.
- Mlíkovský, J. (2003). Brown Fish Owl (Bubo zeylonensis) in Europe: past distribution and taxonomic status. pg. 61-65
- Guerra Rodríguez, Carmen. "Avifauna del pleistoceno superior-holoceno de las Pitiusas: passeriformes y sus depredadores." (2015). Unpublished.
- BirdLife International (2016). "Pinguinus impennis". IUCN Red List of Threatened Species. 2016: e.T22694856A93472944. doi:10.2305/IUCN.UK.2016-3.RLTS.T22694856A93472944.en. Retrieved 19 November 2021.
- Robischon, Marcel (February 2015). "Blue Tigers, Black Tapirs, & the Pied Raven of the Faroe Islands: Teaching Genetic Drift Using Real-Life Animal Examples". The American Biology Teacher. 77 (2): 108–112. doi:10.1525/abt.2015.77.2.5. JSTOR 10.1525/abt.2015.77.2.5. S2CID 85886338.
- BirdLife International (2018). "Numenius tenuirostris". IUCN Red List of Threatened Species. 2018: e.T22693185A131111201. doi:10.2305/IUCN.UK.2018-2.RLTS.T22693185A131111201.en. Retrieved 13 November 2021.
- García, E. & Patterson, A. (2020) Where to watch birds in southern and western Spain. Bloomsbury Publishing, 400 pages.
- BirdLife International (2018). "Geronticus eremita". IUCN Red List of Threatened Species. 2018: e.T22697488A130895601. doi:10.2305/IUCN.UK.2018-2.RLTS.T22697488A130895601.en. Retrieved 19 November 2021.
- Andalusian Buttonquail declared extinct in Spain
- Salvador, A. (2009). Lagartija balear–Podarcis lilfordi (Günther, 1874). Enciclopedia Virtual de los Vertebrados Españoles. Madrid, Spain: Museo Nacional de Ciencias Naturales. http://www. vertebradosibericos. org/(10 May 2018).
- Day, D. (1989). Vanished species. Popular Culture Ink.
- Crivelli, A.J. (2006). "Chondrostoma scodrense". IUCN Red List of Threatened Species. 2006: e.T61345A12465545. doi:10.2305/IUCN.UK.2006.RLTS.T61345A12465545.en. Retrieved 13 January 2018.
- Freyhof, J. & Kottelat, M. (2008). "Coregonus bezola". IUCN Red List of Threatened Species. 2008: e.T135556A4144562. doi:10.2305/IUCN.UK.2008.RLTS.T135556A4144562.en. Retrieved 13 January 2018.
- Freyhof, J. & Kottelat, M. (2008). "Coregonus fera". IUCN Red List of Threatened Species. 2008: e.T135627A4165119. doi:10.2305/IUCN.UK.2008.RLTS.T135627A4165119.en. Retrieved 13 January 2018.
- Freyhof, J. & Kottelat, M. (2008). "Coregonus gutturosus". IUCN Red List of Threatened Species. 2008: e.T135506A4134620. doi:10.2305/IUCN.UK.2008.RLTS.T135506A4134620.en. Retrieved 13 January 2018.
- Freyhof, J. & Kottelat, M. (2008). "Coregonus hiemalis". IUCN Red List of Threatened Species. 2008: e.T135671A4175929. doi:10.2305/IUCN.UK.2008.RLTS.T135671A4175929.en. Retrieved 13 January 2018.
- Freyhof, J. & Kottelat, M. (2008). "Coregonus oxyrinchus". IUCN Red List of Threatened Species. 2008: e.T5380A11126034. doi:10.2305/IUCN.UK.2008.RLTS.T5380A11126034.en. Retrieved 13 January 2018.
- Freyhof, J. & Kottelat, M. (2008). "Coregonus restrictus". IUCN Red List of Threatened Species. 2008: e.T135570A4149314. doi:10.2305/IUCN.UK.2008.RLTS.T135570A4149314.en. Retrieved 13 January 2018.
- Freyhof, J.; Kottelat, M. (2008). "Eudontomyzon sp. nov. 'migratory'". IUCN Red List of Threatened Species. 2008: e.T135505A4134478. doi:10.2305/IUCN.UK.2008.RLTS.T135505A4134478.en. Retrieved 13 November 2021.
- Freyhof, J. & Kottelat, M. (2008). "Gasterosteus crenobiontus". IUCN Red List of Threatened Species. 2008: e.T135637A4167779. doi:10.2305/IUCN.UK.2008.RLTS.T135637A4167779.en. Retrieved 13 January 2018.
- Freyhof, J. & Kottelat, M. (2008). "Romanogobio antipai". IUCN Red List of Threatened Species. 2008: e.T135636A4167651. doi:10.2305/IUCN.UK.2008.RLTS.T135636A4167651.en. Retrieved 13 January 2018.
- Freyhof, J. & Kottelat, M. (2008). "Salvelinus neocomensis". IUCN Red List of Threatened Species. 2008: e.T135421A4127253. doi:10.2305/IUCN.UK.2008.RLTS.T135421A4127253.en. Retrieved 13 January 2018.
- Freyhof, J.; Kottelat, M. (2008). "Stenodus leucichthys". IUCN Red List of Threatened Species. 2008: e.T20745A9229071. doi:10.2305/IUCN.UK.2008.RLTS.T20745A9229071.en. Retrieved 19 November 2021.
- World Conservation Monitoring Centre (1996). "Siettitia balsetensis". IUCN Red List of Threatened Species. 1996: e.T20207A9179037. doi:10.2305/IUCN.UK.1996.RLTS.T20207A9179037.en. Retrieved 13 January 2018.
- Malicky, H. (2014). "Hydropsyche tobiasi". IUCN Red List of Threatened Species. 2014: e.T10332A21426347. doi:10.2305/IUCN.UK.2014-1.RLTS.T10332A21426347.en. Retrieved 13 November 2021.
- Extinction and Hope
- British Wildlife Vol. 11 (1999). British Wildlife Pub.
- "IUCN Red List of Threatened Species: Pseudoyersinia brevipennis". IUCN Red List of Threatened Species. 19 December 2019.
- Martínez, C. (2019) Un piojo que podría extinguirse con el lince ibérico. MNCN-CSIC.
- Jackson, Angus. "Ivell's Sea Anemone". MarLIN - The Marine Life Information Network. The Marine Biological Association of the UK. Retrieved 8 June 2022.
- Triantis, K. (2017). "Zonites siphnicus". IUCN Red List of Threatened Species. 2017: e.T171588A85579865. doi:10.2305/IUCN.UK.2017-3.RLTS.T171588A85579865.en. Retrieved 20 November 2021.
- IUCN
- Albrecht, C.; Hauffe, T.; Reischütz, P. (2011). "Graecoanatolica macedonica". IUCN Red List of Threatened Species. 2011: e.T41027A10390353. doi:10.2305/IUCN.UK.2011-1.RLTS.T41027A10390353.en. Retrieved 12 November 2021.
- Hauffe, T.; Albrecht, C.; Schreiber, K.; Seddon, M.B. (2010). "Ohridohauffenia drimica". IUCN Red List of Threatened Species. 2010: e.T15187A4500356. doi:10.2305/IUCN.UK.2010-4.RLTS.T15187A4500356.en. Retrieved 13 November 2021.
- Prie, V. (2010). "Belgrandia varica". IUCN Red List of Threatened Species. 2010: e.T155668A4818436. doi:10.2305/IUCN.UK.2010-4.RLTS.T155668A4818436.en. Retrieved 13 November 2021.
- IUCN
- Martínez–Ortí, A. L. B. E. R. T. O., & Borreda, V. (2012). New systematics of Parmacellidae P. Fischer 1856 (Gastropoda, Pulmonata), with the recovery of the genus–name Drusia Gray 1855 and the description of Escutiella subgen. nov. Journal of Conchology, 41(1), 1-18.
- Triantis, K. (2017). "Zonites embolium. The IUN Red List of Threatened Species 2017: e.T171211A85578264". IUCN Red List of Threatened Species. 2017. doi:10.2305/IUCN.UK.2017-3.RLTS.T171211A85578264.en. Retrieved 26 May 2022.


















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